Step0
Initial state
Total abundance0
Across all occupied patches
Occupied cells0
Fraction of available habitat
Front radius0
Maximum occupied hex distance
Mean trait φ0
Population‑weighted
Adaptation debt0
Mean |φ − θ|
Trait diversity0
Shannon‑normalized
Patchiness0
Occupied‑neighbor edge fraction
Population relative to local carrying capacity
high
0
Idle

Demography and spread

Evolutionary tracking

Trait distribution

Radial profile

Fluxes